heterogeneity

There are many examples in the literature of a positive correlation between the distribution of a species and its local abundance, i.e., widely occurring species tend to be more abundant locally when they do occur. Such relations have been documented over a wide range of taxa and spatial scales. There are five major hypotheses seeking to explain the relation: Random placement, Sampling error, Niche width, Demography, and Metapopulation dynamics. However, there is little evidence to distinguish between them, especially for plants.

The intermediate disturbance hypothesis has been the focus of considerable analysis in terrestrial and aquatic systems. This model predicts that species diversity will be highest at intermediate frequencies of disturbance. Despite numerous theoretical and empirical analyses, the utility of the model is still the subject of intense debate.